How can mutations over millions of years lead to new species?

Mutations lead to all sorts of variation in organisms and natural selection selects the best able to adapt to the immediate environment by becoming reproductively successful. Thus alleles change over time in populations of organisms and evolution occurs. One; anagenesis. Where a species just evolves so much that it becomes another species. ( rare and controversial ) Cladogenesis. Where species are split asunder by physical or behavioral reasons and split into two different species.