Who is harmed in a thorn acacia and acacia ant relationship?

In the mutualistic relationship between thorn acacia trees and acacia ants, the acacia tree provides shelter and food (nectar and protein-rich Beltian bodies) to the ants, while the ants protect the tree from herbivores and competing plants. However, if the relationship becomes imbalanced, the acacia tree can be harmed if the ants become overly aggressive, attacking beneficial insects or even larger organisms that do not threaten the tree. Additionally, if the tree's growth is stunted or compromised, it may not produce enough resources for the ants, leading to a decline in both populations.