In a 1999 Toyota 4Runner, the fuse for the air conditioning (AC) and rear defogger is located in the fuse box under the dashboard on the driver's side. Specifically, you should look for the fuse labeled "AC" for the air conditioning and "Rear Defog" for the rear defogger. Always refer to the owner's manual for the exact fuse numbers and locations, as configurations can vary. If a fuse is blown, it will need to be replaced to restore functionality.
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