This is unlikely to ever be definitively known, as the fossil record of early tetrapods is incomplete, and the vast gulf of time that separates us from them (360+ million years) compounds the problem. However, we do have some fossils, and from them it is clear that the earliest tetrapods had more than 5 toes on each foot (they were polydactylous). Acanthostega had 8, Ichthyostega had 7 and Tulerpeton had 6 - there seems to have been a trend of reduction in the number of toes over time in the tetrapod lineage. The early Anthracosaurs seem to have had 5 toes ancestrally, a trait which they might have passed on to their amniote descendants. However, it seems as though even the early tetrapods like Acanthostega were effectively five-toed, as their forward toes were combined into a single flesh-covered digit.
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