How are quill feathers adapted to their functions?

Quill feathers, or contour feathers, are primarily adapted for flight and insulation. Their rigid structure provides the necessary aerodynamic shape for lift and maneuverability, while their lightweight design minimizes drag. Additionally, quill feathers have a central shaft with barbs that interlock to create a smooth surface, enhancing aerodynamic efficiency. The arrangement of these feathers also helps in maintaining body temperature by providing insulation against the elements.