Where did the Long-Evans Hooded rat originate?

Long-Evans Hooded rats are an outbred strain from several female Wistar rats and a wild gray Norway rat by Drs. Long and Evans in 1915. Wistar rats are themselves an outbred strain of albino rats of the species "Rattus norvegicus" or Norway rat developed at th Wistar Institute in Philadelphia in 1906 for biological and medical use.