What conclusions did the archaeologist konrad spindler make about the frozen man found in the alps?

Otzi the Iceman was found in 1991. Based on the axe found with the mummy, he dated it to about 4000 years old. Otzi was about 45 years old, stood about 5'5" and weighed about 110 pounds. His last meal included Ibex meat and wheat grains. Copper and arsenic in his hair led to a supposition that he had previously worked in copper smelting. Later, he was probably a shepherd in the mountains, based on wear of his tibia, femur and pelvis. His clothing and tools were intact, leading to conclusions about his lifestyle and status. DNA showed he is closely related to southern europeans, including 19 present day Tyrolean men. One paper states that he had Neanderthal DNA.