The exine, the outer layer of the pollen grain, should be hard to protect the delicate genetic material inside from environmental stresses, such as UV radiation, desiccation, and microbial attack. A tough exine ensures that the pollen can survive the journey to the stigma of a compatible flower, which is crucial for successful fertilization and reproduction. Additionally, a hardened exine aids in the dispersal of pollen by wind or animals, ensuring that it reaches appropriate locations for germination.
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