Genetic and environmental.
Genetic is "hard coded"into the DNA and RNA of the plant or animal, where as environmental is the physical expression of the genetic - this may vary according to environmental conditions and pressures.
A simple example is that a Ficus tree will have large, dark green leaves when in its natural semi-tropical environment, however if the same plant was to be transported to a hot, dry area the leaves would eventually decrease in size and colour in order to reduce the amount of water lost by the plant (and possibly sun burn)
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